Two new species, Mojavemys galushai and Phelosaccomys neomexicanus are desc
ribed from the Barstovian of New Mexico. Both of these species are assigned
to a new subfamily, the Mojavemyinae. It is uncertain as to whether this n
ew subfamily should be referred to the Heteromyidae or Geomyidae.
Both Mojavemys an Phelosaccomys retain some heteromyid characteristics whil
e developing advanced traits of geomyids. It is possible that these genera
represent transitional forms between a heteromyid ancestor and advanced geo
myines, thus deriving the Geomyidae directly from the Heteromyidae sometime
in the middle Tertiary. It is also possible that the Mojavemyinae are a su
bfamily of heteromyids only convergent with geomyids. A third possibility i
s that the mojavemyines are among a group of middle Tertiary geomyoids from
which both Recent heteromyids and geomyids were derived.