The Cenozoic in southern Australia contains many foraminifera endemic
to the region in neritic (intermediate- to shallow-water) facies. They
were mostly epifaunal and inhabited waters to some 300 m deep. This e
ndemism is first obvious in the later Eocene when Maslinella, Crespini
na and Wadella, among others, evolved. More than half of the Eocene en
demic species disappeared in the Eocene or Oligocene. There followed i
n the Oligocene the evolution of such species as Parrellina imperatrix
and Astrononion centroplax. The Miocene was a time of slightly reduce
d endemism and is characterized by migration into the region of many l
arger (sub)tropical taxa such as Lepidocyclina and Cycloclypeus. The l
ong-ranging genus Notorotalia emerged about 50 Ma ago and is still com
mon in modern southern mid-latitude waters. The youngest common extant
species which made their first appearance in the Pliocene-Quaternary
include Dircorbis dimidiatus and Parredicta porifera, both with a test
up to 1.5 mm in diameter. A similar pattern has been recorded in New
Zealand. Four phases of endemism can be recognized: later Eocene, Olig
ocene, Miocene and Pliocene-Quaternary. It appears that the four phase
s were all stimulated in response to major marine transgressions, resp
ectively the Wilson Bluff (=Khirthar), Aldingan, Clifton-Longfordian a
nd Hallet Cove-Glanville transgressions. Probably they signal four imp
ortant stages in the transformation of water masses along the southern
continental margin.