Two unusual Chironomini (Diptera : Chironomidae) from Australian rainforest streams: One new genus and a neotropical genus new for the region

Two genera of chironomine Chironomidae are reported: Anuncotendipes, gen, n ., type species Anuncotendipes anstralotropicus sp. n. and included Anuncot endipes kakadu sp. n., and Nandeva Wiedenbrug, Reiss and Fittkau as new to Australia for the species Nandeva fittkaui sp. n. The pupa and adults are k nown for each genus, but the larvae only for Anuncotendipes. The female of Nandeva is described for the first time. Pupal exuviae of both genera occur in smaller streams in the wet tropics of Far North Queensland, acid A. kak adu is lotic in the Northern Territory. Phylogenetic analysis of the subfam ily Chironominae at generic level suggests that Anuncotendipes is a sister genus to Rheomus in the Harnischia-complex of genera. Nandeva shows converg ent resemblance to the tribe Tanytarsini, but more likely belongs to an aus tral radiation of Chironomini including Nilodosis Keiffer, Fissimentum Cran ston and Nolte and Imparipecten Freeman. These genera show tropical Gondwan an tracks linking Australia with the Afrotropics and Neotropics.