We report tree seedling (mostly Picea engelmannii, some Abies lasiocarpa, v
ery infrequent Pinus contorta) invasion into meadows at upper timberline in
the Snowy Range, Wyoming, from 1994 to 1996. We used gradient analysis to
relate this to environmental patterns, particularly plant community structu
re (as aggregates of plant life-forms) and persistence of snowpack in 1995
and 1996. Tree seedlings established best at sites where snow melted earlie
st; the site with the shortest growing season had fewest seedlings. Microsi
tes dominated by vascular plants admitted few or no seedlings; seedlings we
re most likely to be found where cryptogams dominated. These findings have
implications for forest advance under some climate scenarios and succession
al hypotheses.