The optimal equilibrium harvesting policy is investigated for an age-d
ependent population of females whose life history consists of two stag
es termed eggs (or juveniles) and adults. Using a continuous-time line
ar model, we consider admissible harvesting policies in which a certai
n fraction of individuals of fixed ages is harvested per unit time in
both stages to bring the population to an equilibrium level. Determina
tion of the harvest rate that maximizes the sustainable yield, subject
to a linear ecological or economic constraint, leads to a nonlinear,
nonconvex optimization problem. The optimal policy is shown to consist
of harvesting at most three ages. Thus, we say that the harvest is tr
imodal. In one stage, at most two ages are harvested, with the oldest
being harvested completely; in the other, at most one age is harvested
completely. In each stage, the age totally removed, if present, is ol
der than the surplus age, which is the age at which the proportion of
the expected number of eggs multiplied by the proportion of the expect
ed number of adults first exceeds one. The three harvesting ages are d
ependent on the birth, maturation, and death rates and on the economic
parameters of the problem. A simple algorithm to find the optimum har
vesting strategy is described. (C) Elsevier Science Inc., 1997.