The age difference between the two main globular cluster subpopulations in
the Virgo giant elliptical galaxy NGC 4472 (M49) has been determined using
Hubble Space Telescope WFPC2 images in the F555W and F814W filters, accurat
e photometry has been obtained for several hundred globular clusters in eac
h of the two main subpopulations, down to more than 1 mag below the turnove
r of their luminosity functions. This allows precise determinations of both
the mean colors and the turnover magnitudes of the two main subpopulations
. By comparing the data with various population synthesis models, the age-m
etallicity pairs that fit both the observed colors and magnitudes have been
identified. The metal-poor and the metal-rich globular clusters are found
to be coeval within the errors (similar to Gyr). If one accepts the validit
y of our assumptions, these errors are dominated by model uncertainties. A
systematic error of up to 4 Gyr could affect this result if the blue and th
e red clusters have significantly different mass distributions. However, th
at one subpopulation is half as old as the other is excluded at the 99% con
fidence level. The different globular cluster populations are assumed to tr
ace the galaxy's major star formation episodes. Consequently, the vast majo
rity of globular clusters (and by implication the majority of stars) in NGC
4472 formed at high redshifts but by two distinct mechanisms or in two epi
sodes. The distance to NGC 4472 is determined to be 15.8 +/- 0.8 Mpc, which
is in excellent agreement with six of the seven Cepheid distances to Virgo
Cluster spiral galaxies. This implies that the spiral and elliptical galax
ies in the main body of Virgo are at the same distance.