M. Cresti et F. Ciampolini, Ultrastructural characteristics of pollen development in Vitis vinifera L-(cv. Sangiovese), VITIS, 38(4), 1999, pp. 141-144
Ultrastructural studies on the development of microspores to ripe pollen (c
v. Sangiovese) showed that the diploid microsporocyte undergoes meiosis in
order to form a tetrad of microspores sticked together by a callose wall. A
fter callose digestion, 4 independent microspores were released. With succe
ssive growth, the single nucleus of each microspore undergoes an asymmetric
al division forming the bicellular pollen grain (vegetative cells containin
g the generative cells). Only after pollination and pollen tube growth the
two sperms will be formed. The mature pollen of Sangiovese is tricolpate; t
he exine is scabrate-verrucate in the equatorial zone and foveolate in the
polar region.