In this paper, we propose a consensus sequence for a putative complete Tira
nt retrotransposon. Several defective copies, as well as relevant sequences
available in databases have been analyzed. The putative complete Tirant el
ement is 8533 bp long, and presents all the structural features of a retrov
iruslike transposable element of the gypsy family. It contains three ORFs (
open reading frames) that encode putative products resembling the retrovira
l Gag, Pol, and Env proteins. Southern blot analyses show that complete and
defective Tirant elements are widespread in Drosophila melanogaster. The d
ifferent hybridization patterns observed in several natural populations of
this species suggest that Tirant is an active element.