Relational color constancy, which refers to the constancy of perceived rela
tions between surface colors under changes in illuminant, may be based on t
he computation of spatial ratios of cone excitations. As this activity need
occur only within rather than between cone pathways, relational color cons
tancy might be assumed to be based on relative luminance processing. This h
ypothesis was tested in a psychophysical experiment in which observers view
ed simulated images of Mondrian patterns undergoing colorimetric changes th
at could be attributed either to an illuminant change or to a nonilluminant
change; the images were isoluminant, achromatic, or unmodified. Observers
reliably discriminated the two types of changes in all three conditions, im
plying that relational color constancy is not based on luminance cues alone
. A computer simulation showed that in these isoluminant and achromatic ima
ges spatial ratios of cone excitations and of combinations of cone excitati
ons were almost invariant under illuminant changes and that discrimination
performance could be predicted from deviations in these ratios. (C) 2000 Op
tical Society of America. [S0740-3232(00)01502-7] OCIS codes: 330.0330, 330
.1720, 330.1690, 330.4060.