Spatial orientation in five species of ants was described by measurements o
f trail length and percentage of trail along structural guidelines (edges,
grooves, crestlines). In the absence of structural guidelines, carpenter an
t trails were essentially straight. Carpenter ants traveled significantly f
aster on structural guidelines than on the ground through vegetation. As am
bient temperature decreased, ants slowed down more quickly when traveling t
hrough vegetation compared with ants traveling on structural guidelines. A
detour experiment indicates that ants save on both time and energy by using
structural guidelines.