We developed the landscape age-class demographics simulator (LADS) to model
historical variability in the amount of old-growth and late-successional f
orest in the Oregon Coast Range over the past 3,000 years. The model simula
ted temporal and spatial patterns of forest fires along with the resulting
fluctuations in the distribution of forest age classes across the landscape
. Parameters describing historical fire regimes were derived from data from
a number of existing dendroecological and paleoecological studies. Our res
ults indicated that the historical age-class distribution was highly variab
le and that variability increased with decreasing landscape size. Simulated
old-growth percentages were generally between 25% and 75% at the province
scale (2,250,000 ha) and never fell below 5%. In comparison, old-growth per
centages varied from 0 to 100%, at the late-successional reserve scale (40,
000 ha). Province-scale estimates of current old-growth (5%) and late-succe
ssional forest (11%) in the Oregon Coast Range were lower than expected und
er the simulated historical fire regime, even when Potential errors irt our
parameter estimates were considered. These uncertainties no, however, limi
t our ability to Precisely define ranges of historical variability. Our res
ults suggest that in areas where historical disturbance regimes were charac
terized by large infrequent fires, management of forest age classes based o
n a range of historical variability may be feasible only at relatively larg
e spatial scales. Comprehensive landscape management strategies will need t
o consider other factors besides the percentage of old forests on the lands
cape, including the spatial pattern of stands and the rates and pathways of
landscape change.