Tetrapods include the only fully terrestrial vertebrates, but they also inc
lude many amphibious, aquatic and flying groups, They occupy the highest le
vels of the food chain on land and in aquatic environments. Tetrapod evolut
ion has generated great interest but the earliest phases of their history a
re poorly understood. Recent studies have questioned long-accepted hypothes
es about the origin of the pentadactyl limb, the phylogeny of tetrapods and
the environment in which the first tetrapods lived.