The evolutionary causes of small clutch sizes in tropical and Southern Hemi
sphere regions are poorly understood. Alexander Skutch proposed 50 years ag
o that higher nest predation in the south constrains the rate at which pare
nt birds can deliver food to young and thereby constrains clutch size by Li
miting the number of young that parents can feed. This hypothesis for expla
ining differences in clutch size and parental behaviors between latitudes h
as remained untested. Here, a detailed study of bird species in Arizona and
Argentina shows that Skutch's hypothesis explains clutch size variation wi
thin North and South America. However, neither Skutch's hypothesis nor two
major alternatives explain differences between latitudes.