Anthropogenic increases in the atmospheric concentration of carbon dio
xide and other greenhouse gases are predicted to cause a warming of th
e global climate by modifying radiative forcing(1). Carbon dioxide con
centration increases may make a further contribution to warming by ind
ucing a physiological response of the global vegetation-a reduced stom
atal conductance, which suppresses transpiration(2). Moreover, a CO2-e
nriched atmosphere and the corresponding change in climate may also al
ter the density of vegetation cover, thus modifying the physical chara
cteristics of the land surface to provide yet another climate feedback
(3-6). But such feedbacks from changes in vegetation structure have no
t yet been incorporated into general circulation model predictions of
future climate change. Here we use a general circulation model iterati
vely coupled to an equilibrium vegetation model to quantify the effect
s of both physiological and structural vegetation feedbacks on a doubl
ed-CO2 climate. On a global scale, changes in vegetation structure are
found to partially offset physiological vegetation-climate feedbacks
in the long term, but overall vegetation feedbacks provide significant
regional-scale effects.