Jms. Costas, A new revision and taxonomic position for the Cantabrian Leptodirinae: Sections Quaestus and Speonomidius (Coleoptera : Cholevidae), ANN SOC ENT, 36(1), 2000, pp. 45-59
A taxonomic review of the Cantabrian Leptodirinae has been carried out. The
basis for this review is the study of certain important external morpholog
ical characteristics, but in particularly, the examination of the male and
female genitalia. As a result, we propose a new position in sections and th
ree new taxonomic categories, Cantabrogeus nov, gen., Amphogeus nov, subgen
., and Samanolla nov. subgen., as well as the "new status" of Speonomidius.