A. Westman et al., The phylogeny of the genus Leucorrhinia and the evolution of larval spines(Anisoptera : Libellulidae), ODONATOLOGI, 29(2), 2000, pp. 129-136
A cladistic analysis of the genus Leucorrhinia, based on adult morphologica
l characters, found one most parsimonous tree with a consistency index of 0
.35. The evolution of large dorsal larval spines was mapped on the resultin
g tree. This mapping suggests that the presence of spines is the primitive
state within Leucorrhinia and that they have disappeared on five different
occasions, or have disappeared twice on lower branches and reappeared three
times higher up in the tree.