The purpose of this paper is to explore relationships among frogs that have
been (variously) placed in the Ranoidea. Phylogenetic analyses were conduc
ted on data sets including 10 morphological characters and sequences from p
ortions of the 12S, 16S, and tRNA(val) mitochondrial genome. The resulting
phylogenetic trees suggest that (1) the sooglossids are the sister group to
all other microhyloid and ranoid frogs, (2) the Microhylidae may be the si
ster group of the Hyperoliidae. (3) the Hyperoliidae as classically defined
may be diphyletic, (4) Hemisus appears to be a microhylid. and (4) the man
telline frogs may be clustered within the Rhacopholidae. Analysis shows tha
t both morphological and molecular characters contain significant phylogene
tic signal. Nonetheless, more morphological and molecular characters will b
e necessary to resolve completely the relationships among the deep branches
of the Ranoidea.