This paper formulates the problem of measuring bilateral symmetry of object
s analytically for landmark data, and develops various new testing procedur
es and exploratory data analyses. The development is linked with the fundam
ental biological problem of measuring directional asymmetry and fluctuating
asymmetry. We distinguish two types of symmetry, object symmetry and match
ing symmetry, and provide tests under assumptions of isotropic landmark var
iability as well as non-isotropy. The tests require novel statistical and g
eometrical analyses of the Procrustes shape manifold. For describing compon
ents of symmetry and asymmetry within samples, an extension of principal co
mponent analysis is introduced and illustrated by appropriate deformation t
echniques. Various real examples are used to illustrate the practical relev
ance of this work.