The spatial scale of variability in small-mammal populations

We studied small-mammal populations across a range of spatial scales to det ermine which scales were relevant to demographic variability. We predicted that a scale of variability in population structure would occur at the scal e of dispersal, which has previously been described as <200 m for some smal l-mammal species. Systematic live-trapping surveys were conducted on nested grids at three scales: 1) extent = 4900 ha, grain = 1000 m; 2) extent = 30 6 ha, grain = 250 m; and 3) extent = 31 ha, grain = 125 m. Prior to the pre sent study, small-mammal populations had not been systematically live-trapp ed across a similar range of scales. From 1996 to 1998, surveys were conduc ted on a landscape intensively managed for timber, and on a reference lands cape. Spatial analysis of abundance data demonstrated that small-mammal pop ulations (four species) exhibited positive spatial autocorrelation at dista nces of between 133 and 533 m depending on the species and the landscape. N o higher-order population structure was detected. Thus, within the range of scales we sampled, variability in small-mammal abundance occurred over sho rt distances (i.e., 133-533 m).