Archeological analysis of faunal remains and of lithic and bone tools has s
uggested that hunting of medium to large mammals was a major element of Nea
nderthal subsistence. Plant foods are almost invisible in the archeological
record, and it is impossible to estimate accurately their dietary importan
ce. However, stable isotope (delta(13)C and delta(15)N) analysis of mammal
bone collagen provides a direct measure of diet and has been applied to two
Neanderthals and various faunal species from Vindija Cave, Croatia. The is
otope evidence overwhelmingly points to the Neanderthals behaving as top-le
vel carnivores, obtaining almost all of their dietary protein from animal s
ources. Earlier Neanderthals in France and Belgium have yielded similar res
ults, and a pattern of European Neanderthal adaptation as carnivores is eme
rging. These data reinforce current taphonomic assessments of associated fa
unal elements and make it unlikely that the Neanderthals were acquiring ani
mal protein principally through scavenging. Instead, these findings portray
them as effective predators.