The richness and coexistence of pteridophyte species were studied at the me
soscale level in the NE Iberian Peninsula (Catalonia and Andorra) using a g
rid system with 10x10 km cell-size. The number of pteridophytes, the number
of monolete and trilete ferns (two spore patterns), and the number of poly
ploid and diploid ferns were studied in 371 of such units. The results sugg
est: (a) a significant relationship of pteridophyte species richness with h
igher-taxon richness (genera, family, order, class); (b) a significant rela
tionship between pteridophyte richness and maximum altitude (positively rel
ated), bedrock type (higher richness in siliceous types), and distance to t
he coast (interacting with altitude); (c) a significant positive relationsh
ip of monolete/trilete ratio with altitude (in the study area) and with lat
itude (at the European scale); (d) a significantly higher polyploid/diploid
ratio for monolete ferns than for trilete ferns, and a significant relatio
n of this ratio with altitude. We conclude that at the scale studied, the d
istribution of pteridophyte richness is mainly related to terrain and clima
te parameters and that the monolete/trilete pattern may be explained by the
higher polyploidy rate and better migration capacity in monolete species.