Some populations of wild chimpanzees (Pan troglodytes) use hammers and
anvils of stone or wood to crack open nuts for food. Others do not. T
he aim of this study was to ask why one non-nut-cracking population, i
n the Lope' Reserve, Gabon, lacks this useful form of tool use. We tes
ted 10 hypotheses: (1) nuts are absent; (2) nuts are few; (3) nuts are
unsuitable; (4) hammers are absent; (5) hammers are unsuitable; (6) a
nvils are absent; (7) anvils are unsuitable; (8) nuts are displaced by
better food items; (9) intelligence is insufficient; and (10) knowled
ge is insufficient. All but the last are clearly falsified, leaving by
exclusion the likelihood that Lope's chimpanzees lack the technology-
knowledge of appropriate technique-to exploit this resource. Thus, the
behavioral differences across populations of these apes are cultural
and not environmentally dictated. This explanation is congruent with t
he distribution of chimpanzee nut-cracking across Africa.