Margachitina margaritana is a distinctive, chain-forming chitinozoan taxon
of the uppermost Telychian Stage (Llandovery Series) to lower Homerian Stag
e (Wenlock Series). M. margaritana is shown to be the end member of a morph
ological lineage that developed from Calpichitina densa via the intermediat
e Margachitina banwyensis sp. nov. A local M. banwyensis Biozone, which cor
relates with the upper lapworthi graptolite Biozone, is proposed for the Ba
nwy River section. The morphological lineage described provides a tool for
the enhanced subdivision of the uppermost Llandovery Series, which may prov
e global in extent. This research suggests also that the base of the Wenloc
k Series may correlate with a level below the centrifugus graptolite Biozon
e and it also brings into question the systematic status of Calpichitina an
d Margachitina, although both genera are retained herein.