Lady beetles (Coleoptera : Coccinellidae) from Chinese hemlocks infested with the hemlock woolly Adelgid, Adelges tsugae Annand (Homoptera : Adelgidae)

Fifty-four species of lady beetles were collected from three Chinese hemloc k species, Tsuga dumosa (D. Don) Eichler, T. forrestii Downie and T. chinen sis (Franchet) Pritzel, in Yunnan, Sichuan and Shaanxi provinces of China a s part of a search for natural enemies of the hemlock woolly adelgid, Adelg es isugae Annand. Twenty new species are described: Clitostethus wenbishanu s Yu, Scymnus (Neopullus) ningshanensis Yu and Yao, S. (N.) lijiangensis Yu , S. (N.) lycotropus Yu, S. (N.) nigromarginalis Yu, S. (Parapullus) tsugae Yu and Yao, S. (Scymnus) unciformis Yu, S. (S.) paracrinitus Yu, S. (Pullu s) nigrobasalis Yu, S. (P.) baoxingensis Yu, S. (P.) gucheng Yu, S. (P.) ge minus Yu and Montgomery, S. (P.) heyuanus Yu, S. (P.) robustibasalis Yu, S. (P.) jaculatorius Yu, Pseudoscymnus heijia Yu and Montgomery, P. ocellatus Yu, Shirozuella quadrimacularis Yu, and Sh. nibagou Yu. New synonyms place d here are: Oenopia yunlongensis Jing for O. deqenensis Jing, and O. pomien sis Jing, O. gonggarensis Jing and O. picithoroxa Jing for O. billieti (Mul sant). Harmonia quadripunctata (Pontoppidan) is a new record for China. Key s are provided to separate the known species of Clitostethus Wiese, Scymnus (Neopullus) Sasaji, and Scymnus (Scymnus) Kugelann in China. Nine species are recorded as predators of the hemlock woolly adelgid: Oenopia billieti ( Mulsant), Adalia conglomerata (L.), Calvia championorum Booth, P. ocellatus , S. (N.) camptodromus Yu et Liu, S. (N.) sinuanodulus Yu ct Yao, S. (N.) n ingshanensis, S. (P.) yunshanpingensis Yu, and S. (P.) geminus.