I. Arenillas et al., The Cretaceous/Paleogene (K/P) boundary at Ain Settara, Tunisia: Sudden catastrophic mass extinction in planktic foraminifera, J FORAMIN R, 30(3), 2000, pp. 202-218
The quantitative study and high resolution sampling of an essentially conti
nuous and expanded Cretaceous/Paleogene (K/P) boundary section in Tunisia a
llow us to test the model of extinction in planktic foraminifera. The extin
ction at the Ain Settara section occurred over a short period of time simil
ar to the Tunisian sections at El Kef and Elles and the Spanish sections at
Agost, Caravaca and Zumaya, At Ain Settara only 3 species disappeared in t
he latest Maastrichtian, 45 became extinct precisely at the Km boundary and
18 disappeared in the earliest Danian. The species that became extinct at
the K/P boundary constitute about 20% of the individuals in the population
larger than 63 microns and 68% of the species, which suddenly became extinc
t in a catastrophic event precisely coinciding with the layer containing ev
idence for an asteroid impact. Most of these species are large, complex and
low latitude deeper to intermediate dwelling forms. This extinction event
is clearly the most important catastrophic mass extinction recorded in the
history of planktic foraminifera. This pattern of extinction is superimpose
d on a controversial gradual pattern of extinction of 21 species that appar
ently began in the latest Maastrichtian and ended in the early Danian, The
Maastrichtian species that seem to become extinct gradually are generally s
mall, cosmopolitan and simple surface dwellers. The catastrophic mass extin
ction of the 45 species coincident with the K/P boundary is compatible with
the effect of the impact of a large asteroid, whereas the gradual extincti
on of 18 species in the basal Danian could also be attributed to the long t
erm disruptive effect of the impact.