The arctic flora is thought to have originated during the Late Tertiary, ap
proximately 3 million years ago. Plant migration routes during colonization
of the Arctic are currently unknown, and uncertainty remains over where ar
ctic plants survived Pleistocene glaciations. A phylogenetic analysis of ch
loroplast DNA variation in the purple saxifrage (Saxifraga oppositifolia) i
ndicates that this plant first occurred in the Arctic in western Beringia b
efore it migrated east and west to achieve a circumpolar distribution. The
geographical distribution of chloroplast DNA variation in the species suppo
rts the hypothesis that, during Pleistocene glaciations, some plant refugia
were Located in the Arctic as well as at more southern Latitudes.