Is starting a new business more difficult in an emerging industry or in a m
ature industry? The density dependent model of organizational ecology maint
ains that the industry's age is irrelevant; the number of firms currently o
ccupying the market niche determines the industry's competitive structure.
Nevertheless, population-level learning predicts historical asymmetry in en
try barriers. Over time, the average fitness of the surviving population me
mbers increases, making market entry more difficult. At the same time, surv
iving organizations become increasingly spread out across the resource spac
e, providing niches that new firms can exploit. Thus, industry-level evolut
ion systematically alters the environment that both existing organizations
and new firms face. I offer a new specification for the founding rate model
that synthesizes ecological and evolutionary perspectives. Tests of this m
odel in the American automobile industry support its merit. (C) 2000 Academ
ic Press.