Demographic transition theory posits that modernization, particularly in th
e form of urban industrialism, fundamentally alters the environmental conte
xt surrounding fertility decision-making, thereby reducing the advantages o
f having children. While fertility research has either questioned the link
between modernization and fertility or attempted to provide the intervening
links between the two, there has been, little theoretical or empirical ref
inement of the macrosocial/contextual principles of the theory. We argue th
at human ecology and evolutionary theory can help respecify and revitalize
demographic transition theory. Our respecification produces a more logical
account of fertility decline that emphasizes the influences of service econ
omies and the social adaptations attendant on ethnic heterogeneity and prei
ndustrial social complexity. Analyses of changes in total fertility rates i
n approximately 60 less-developed countries suggest: (I) general (if partia
lly ambiguous) support for demographic transition theory, (2) confirmation
of a robust effect of service-sector dominance on fertility, and (3) the im
portance of ethnic homogeneity and preindustrial social complexity to demog
raphic transition.