The actualistic method was applied to assess the causes of asymmetry in lat
itudinal zoning of Devonian, Carboniferous, and Permian faunas, Early Permi
an benthic fauna located northeast of the Pangaea-2 supercontinent is chara
cterized as more cold-water fauna than that located northwest of the contin
ent. This can be explained by the opposing directions of the coastal curren
ts of the northern subpolar gyre at the same latitudes. For the first time,
paleogeographic elements of the fauna at the species level are distinguish
ed in the same manner as is done in modern marine biogeography. A model of
biochore evolution in the Late Paleozoic and Triassic seas is presented.