Ac. Sandford, Trilobite faunas and palaeoenvironmental setting of the Silurian (early Ludlow) Melbourne Formation, central Victoria, ALCHERINGA, 24(3-4), 2000, pp. 153-206
Trilobites are common faunal elements in the Melbourne Formation, a unit of
early Ludlow (upper nilssoni Biozone) age, which crops out extensively in
the Darraweit Guim Province of the Melbourne Zone, central Victoria. New di
agnoses are given for species previously described, including Maurotarion e
uryceps (McCoy, 1876; = Cyphaspis spryi Gregory, 1901), Raphiophorus jikaen
sis (Chapman, 1912; = Ampyx yarraensis Chapman, 1912), Cromus simpliciculus
(Talent, 1964), Cromus spryi (Chapman, 1912), Sthenarocalymene kilmorensis
(Gill, 1945; = Gravicalymene hetera Gill, 1945) and Trimerus harrisoni (Mc
Coy, 1876). A new phacopid genus, Orygmatos is described, represented by th
e species O. yanyeani gen. et sp. nov. Other species newly described includ
e Cromus melbournensis sp. nov., Arcticalymene australis sp. nov., "Ananasp
is" woiwurrungi sp. nov. and Kettneraspis hollowayi sp. nov.
Species composition of the trilobite fauna varies spatially, and a number o
f distinct assemblages can be defined. Abundant trilobite moult configurati
ons are conclusive for interpretation of the benthic fauna as autochthonous
, inferring depth estimations based on benthic community distribution to be
valid. A depth-related succession of communities is recognised and indicat
e the Melbourne Formation was deposited at relatively shallow depths on a b
road, eastwardly deepening shelf, with deposition dominated by storm proces
ses. The palaeoenvironment comprised a BA-I community including the Arctica
lymene australis trilobite assemblage, restricted to very shallow depths (s
imilar to 20 m) on the SW coastal margin of the shelf and preserved in prox
imal tempestite lithologies; and a BA-5 community group containing three di
stinct trilobite assemblages dominated by species of Cromus and a deeper wa
ter fauna, preserved in distal tempestite lithologies and ranging widely ov
er the shelf at depths in the range of maximum storm wave base (similar to
50 - 80 m).