One of the most ubiquitous phenomena of all natural populations is their va
riability in numbers in space and time. However, there are notable differen
ces among populations in the way the population size fluctuates. One of the
major challenges in population and community ecology is to explain and und
erstand this variety and to find possible underlying rules that might be mo
dified from case-to-case. Population variability also has a spatial compone
nt because fluctuations are often synchronized over relatively large distan
ces. Recently, this has led to growing interest in how 'internal' (density-
dependent) processes interact with 'external' factors such as environmental
variability.