Using accurate distances to individual Virgo Cluster galaxies obtained by t
he method of surface brightness fluctuations, we show that Virgo's brightes
t elliptical galaxies have a remarkably collinear arrangement in three dime
nsions. This axis, which is inclined by similar to 10 degrees -15 degrees f
rom the line of sight, can be traced to even larger scales where it appears
to join a filamentary bridge of galaxies connecting Virgo to the rich clus
ter Abell 1367. The orientations of individual Virgo elliptical galaxies al
so show some tendency to be aligned with the cluster axis, as does the jet
of the supergiant elliptical galaxy M87. These results suggest that the for
mation of the Virgo Cluster, and its brightest member galaxies, have been d
riven by infall of material along the Virgo-A1367 filament.