'Lydekkerina' putterilli Broom from the Lystrosaurus Assemblage Zone (Middl
e Beaufort Group, South Africa) is shown to be a paedomorphic rhinesuchid (
Broomistega putterilli gen. nov.) rather than a lydekkerinid or juvenile st
age of the rhinesuchid Uranocentrodon, as previously presumed. The most con
spicuous characters, not related to its paedomorphic condition, include the
shape of the parietals and the structure of the parasphenoid body and exoc
cipitals. The pattern of cranial growth changes in Broomistega has been inv
estigated by comparison of the holotype with two hitherto undescribed immat
ure specimens. For several characters referring to the size and position of
the orbits, and re-modelling of the jugal proportions, retarded timing of
growth transformations, compared to ontogenetic patterns in Permian rhinesu
chids (typified by Muchocephalus) is demonstrated. Some features of the evo
lution of the basicranial region of the skull, in the transition from rhine
suchids to more advanced capitosauroids, are re-evaluated.