We summarize the influences of harvester ants of the genus Pogonomyrmex on
communities and ecosystems. Because of nest densities, the longevity of nes
ts, and the amount of seed harvested and soil handled, harvester ants have
significant direct and indirect effects on community structure and ecosyste
m functioning. Harvester ants change plant species composition and diversit
y near their nests. These changes result from differential seed predation b
y the ants, their actions as seed dispersers and competitors with other gra
nivores, and the favorable soil conditions they create through their diggin
g. Their nest building creates islands of increased nutrient density. In so
me areas, the effects of their activities may be so pervasive that plant co
mmunity structure is strongly influenced. Ant removal studies, which would
reveal their total impact, have generally not been done. Granivore removals
have been conducted in North America where ants are of lesser importance t
han small mammals, in contrast to other areas (except Israel) where ants ar
e dominant granivores. We review the influence of harvester ants on their c
ompetitors, predators, and nest associates, and catalog the factors that in
fluence their foraging patterns and consequently their local distribution.