Fossil plankton an difficult to identify but have formed a major component
of most marine ecosystems throughout geological time. The earliest fossil h
eterotrophs include planktic forms, and subsequent adaptive innovations qui
ckly appeared in the plankton; these include metazoans and animals with har
d parts. Movement into the plankton occurred sporadically throughout geolog
ical history and seems to have been independent of any biological or enviro
nmental forcing mechanism. Subsequent radiations and extinctions in the coh
ort of plankton closely reflect events in the benthos. The diversity of zoo
plankton rose quickly during the early Paleozoic era, but low plankton dive
rsity characterized the late Paleozoic. Significant radiations during early
Mesozoic times led to an overall increase in diversity through the Phanero
zoic eon. As the composition of the zooplankton has changed, so has their e
ffect on biogeochemical cycles.