Genypterus valdesensis. sp. nov. from the Puerto Madryn Formation in Penins
ula Valdes, Argentina is the first fossil ophidiid reported from South Amer
ica and represents the first well preserved record of the genus worldwide.
The specimen consists of a nearly complete articulated skull and some disar
ticulated postcranial elements. A thorough description is given of the oste
ological features of the species, which can be used to recognize it as a di
stinct taxon. The fossil species is characterized by the presence of a fora
men on infraorbital 3 and by the square-shape of infraorbital 4. An array o
f morphological features taken in combination contribute to further differe
ntiate G. valdesensis from the Recent species. This specimen also provides
unequivocal evidence of the presence of the Ophidiiformes in the southweste
rn Atlantic Ocean 14 MA.