Ascidians are urochordates, marine invertebrates with non-feeding motile ch
ordate tadpole larvae, except in the family Molgulidae. Urodele, or tailed,
Molgulids have typical ascidian char-date tadpole larvae possessing fails
with muscle cells, a notochord, and a dorsal hollow nerve cord. In contrast
, anural (or tail-less) Molgulids lack a ta ii and defining chordate featur
es. Molecular phylogenies generated with 18S and 28S ribosomal sequences in
dicate that Molgulid species fall into at least four distinct clades, three
of which have multiple anural members. This refined and expanded phylogeny
allows careful examination of the factors that may have influenced the evo
lution of tail-less ascidians.