The doliolid heart is briefly described: despite the absence of closed
vessels, it produces an efficient circulation alone defined paths wit
hin the haemocoel. When it reverses, the now follows the same paths in
reverse. There are very few cells within the haemocoel fluid, but the
movements of the occasional cell permit entry velocity to the heart t
o be measured, and intracardiac pressures of some 2Pa to be calculated
. Heart volume is around 0.15% of that of the haemocoel.