Based on specimens from Australia and Iran, five species of rhipidognathid
conodonts, Appalachignathus delicatulus Bergstrom, Carnes. Ethington. Votaw
, and Wigley, 1974. Bergstroemognathus extensus (Graves and Ellison, 1941),
B. hubeiensis An (MS) in An, Chen. and Li. 1981. B. kirki Stair and Druce.
1993. and Rhipidognathus? vichangensis (Ni, 1981). are described and revis
ed in terms of multielement morphology. All three genera comprising the Rhi
pidopnathidae are interpreted as having a septimembrate apparatus. partiall
y confirmed by bedding plane assemblages of B. extensus from Victoria. Occu
rrence of A. delicatulus in allochthonous limestones (about the Middle-Uppe
r Ordovician boundary) of central New South Wales is the first record of th
e species outside North America. Recognition of Rhipidognathus? yichangensi
s in Early Ordovician strata of the Canning Basin, reinforces biogeographic
affinities of Australia and South China. The three described species of Be
rgstroemognathus are mainly restricted to late Early Ordovician strata. Ber
gstroemognathus extensus is widely distributed in North America, western Ar
gentina (Precordillera), China, and Australia. Bergstroemognathus hubeiensi
s, described from east-central Iran, has been previously recorded only from
China, while the slightly younger B. kirki seems endemic to central and no
rthern Australia. where it was restricted to shallow, warm water environmen
ts. In contrast. B. extensus and B. hubeiensis inhabited a spectrum of wate
r depths from shallow to deep.