Earth is over 4,500 million years old. Massive bombardment of the planet to
ok place for the first 500-700 million years, and the largest impacts would
have been capable of sterilizing the planet. Probably until 4,000 million
years ago or later, occasional impacts might have heated the ocean over 100
degreesC. Life on Earth dates from before about 3,800 million years ago, a
nd is likely to have gone through one or more hot-ocean 'bottlenecks'. Only
hyperthermophiles (organisms optimally living in water at 80-110 degreesC)
would have survived. It is possible that early life diversified near hydro
thermal vents, but hypotheses that life first occupied other pre-bottleneck
habitats are tenable (including transfer from Mars on ejecta from impacts
there). Early hyperthermophile life, probably near hydrothermal systems, ma
y have been non-photosynthetic, and many housekeeping proteins and biochemi
cal processes may have an original hydrothermal heritage. The development o
f anoxygenic and then oxygenic photosynthesis would have allowed life to es
cape the hydrothermal setting. By about 3,500 million years ago, most of th
e principal biochemical pathways that sustain the modern biosphere had evol
ved, and were global in scope.