Two types of central projections originating from the lateral ocelli h
ave been identified in Lithobius: First, shore retinula axons terminat
e within the first optic neuropil or lamina. They possess varicose swe
llings throughout the depth of the neuropil. Second, long axons pass t
hrough the first neuropil and terminate within the medulla, where up t
o five long collaterals are found that diverge to various synaptic sit
es. A comparison with other Arthropoda suggests that the presence of b
oth short and long retinula axons might be a common feature of the man
dibulates' lateral visual system, while divergence of the long visual
fibers appears to be a unique myriapodan character.