The machaeridian Lepidocoleus sarlei Clarke, 1896, from the Rochester Shale (Silurian) of New York State

Eight new specimens and the holotype of Lepidocoleus sarlei Clarke, 1896 fr om Lagerstatten deposits within the Rochester Shale (Silurian, Wenlock) of New York are described to elucidate the morphology, function and constructi on of the scleritome. The material exhibits an unusual type of dorsal hinge with a pronounced anteroposterior displacement between the two sides. Two specimens have an aberrant sclerite on the right side of segment eleven. On e tightly enrolled specimen may demonstrate a method of defence with the st ronger protected dorsal side oriented outwards. The presence of a segmented body plan is suggested, based on requirements for a functional spiral enro lment. Distinct anterior and posterior morphologies together with the patte rn of rugae provide valuable help in reconstructing disarticulated lepidoco leids.