Z. Kvacek et B. Erdei, Putative proteaceous elements of the Lomatites-type reinterpreted as new Berberis of the European Tertiary, PLANT SYS E, 226(1-2), 2001, pp. 1-12
A characteristic xeromorphic foliage occurring in the European Tertiary was
originally identified as a Proteaceae according to gross morphology, and c
ompared with Grevillea L. and Lomatia L. (Grevillea kymeana Unger. Lomatite
s aquensis Saporta). On the basis of venation patterns and phyllotaxy, thes
e elements are now reinterpreted as extinct species of Berberis L. (Berberi
daceae): Berberis andreanszkyi spec, nov. occurring in the Upper Oligocene
(Eger - Wind brickyard)? and the Middle Miocene (Erdobenye) of Hungary, and
Berberis kymeana (Unger) comb. nov. (syn. Lomatites aquensis Saporta) know
n from the Lower Miocene of Greece (Kymi) and the Oligocene of France (Aix-
en-Provence. Manosque/Bois d'Asson. Cerestre, Bonieux). The associated vege
tation reflects warm-temperate to subtropical subhumid climatic conditions.
Extant barberries with similar foliage occur in the Himalayas and China to
day.