unusual trochiform gastropod, Semizona bella gen. et sp. nov., is described
from the Boda Limestone carbonate mounds (upper Ordovician, Ashgill) of ce
ntral Sweden. A second species, S. glindmeyeri (Rohr, 1996), is recognised
from the Ordovician (Whiterockian) of Nevada. The shell shape and the stron
gly prosocline tangential aperture of Semizona suggest that balancing of th
e shell on the head-foot mass was accomplished by tilting of the axis of co
iling of the shell to about 65 degrees with 10-30 degrees of regulatory det
orsion. The rounded aperture allowed straight contraction of the retractor
muscles, suggesting clamping behaviour often associated with a sedate, graz
ing snail. This agrees with the environmental setting, which suggests a har
d substrate with rich microbial growth. Besides clamping, the subsutural no
des and thick shell were probably effective against predation; repaired inj
uries indicate failed predatory attacks. Semizona shows morphological simil
arities with some pleurotomariin vetigastropods, and with the family Pseudo
phoridae Miller, 1889.