Changes in the abundance of species-especially those that influence wa
ter and nutrient dynamics, trophic interactions, or disturbance regime
-affect the structure and functioning of ecosystems. Diversity is also
functionally important, both because it increases the probability of
including species that have strong ecosystem effects and because it ca
n increase the efficiency of resource use. Differences in environmenta
l sensitivity among functionally similar species give stability to eco
system processes, whereas differences in sensitivity among functionall
y different species make ecosystems more vulnerable to change. Current
global environmental changes that affect species composition and dive
rsity are therefore profoundly altering the functioning of the biosphe
re.