Experimental discovery of the dilaton would provide strong evidence for str
ing theory. A light dilaton could show up in current tests of the inverse s
quare law for gravity at sub-millimeter distances. In the large extra dimen
sion scenario, Kaluza-Klein excitations of the dilaton can also contribute
to the cooling of supernovae. In order to quantify these effects we compute
the couplings of a low energy dilaton to matter. These predominantly arise
from the fundamental dilaton coupling to the gluon field strength, which r
eceives a sizable enhancement from QCD scaling. We show that detection of t
he dilaton will give a direct measurement of the QCD coupling constant at t
he string scale. Particular attention is paid to the size of equivalence pr
inciple violating dilaton couplings.