Stomach contents of northern bottlenose whales Hyperoodon ampullatus stranded in the North Sea

This paper presents information on the stomach contents of four northern bo ttlenose whales Hyperoodon ampullatus (Odontoceti: Ziphiidae) from the nort h-east Atlantic, an area for which there art: few recent data on the feedin g ecology of this species. Two of these whales were relatively recent stran dings, a female stranded in August 1993 at Hargen (the Netherlands) and a m ale stranded in February 1997 on the island of Tasinge (Denmark). Stomach c ontent samples were also examined from a juvenile male stranded in November 1885 at Dunbar (Scotland) and a female stranded in August 1956 an the isla nd of Texel (the Netherlands). Food remains from the four samples consisted almost entirely of cephalopod beaks. Some fish remains were also found in the stomach of the Hargen and T asinge whales, and the latter also had crustacean remains in the stomach. T he cephalopod prey consisted mainly of oceanic cephalopods: Gonatus sp. (pr obably G: fabricii, Cephalopoda: Teuthoidea)? Taonius pavo and Histioteuthi s sp. for the Dunbar whale; Gonatus and Teuthowenia megalops for the Texel whale; Gonatus for the Hargen whale and Gonatus, T. megalops and Taonius pa vo for the Tasinge whale. Other prey species found in the Tasinge specimen included the squid Histioteuthis reversa, H. arcturi, and the octopods Vamp iroteuthis infernalis and Vitreledonella richardi. Based on the size of the lower beaks, the squid eaten included juvenile and mature individuals of t he most important species (Gonatus and Teuthowenia megalops). The fish rema ins consisted of vertebrae of Gadidae and fish eve lenses (Hargen whale) an d two Trisopterus otoliths (Tasinge whale). The results from this study are in agreement with those of previous authors in that cephalopods in general, and G. fabricii in particular, are the mai n prey of the northern bottlenose whale and other toothed whales in norther n latitudes.