Field studies have shown that resistance to convective gas-flow within the
culm-rhizome gas-space system of Phragmites tends to be higher in 'die-back
' as opposed to healthy stands. However, the collection of reliable data an
d its interpretation is complicated by many factors. To help interpret fiel
d measurements and further our understanding of convective flows in general
, we have developed mathematical models based on the humidity-induced conve
ctive-flow generating potential of culms. Among other things, these make it
possible to study the effects on pressure flows of increasing flow resista
nces in rhizomes and between culms, of different pressure-generating potent
ials of interlinked culms and of different numbers of efflux culms. A multi
-culm and rhizome model is described together with some simple examples of
the way it can prove helpful in interpreting some of the field observations
from healthy and die-back sites.
Increased venting resistance reduced flows curvilinearly; increasing the co
unterpressure to mimic those induced by other interlinked living culms redu
ced flows linearly. In some of the multi-culm examples shown counter-pressu
re exerted an effect approximately the same as that of rhizome and venting
resistance, but as culm numbers declined it assumed even greater importance
. The value of conductivity derived from applying pressures to the stubble
of excised culms and measuring the flows induced, proved to be a composite
measure of the effective conductivity of the whole rhizome-culm train rathe
r than that of the rhizome plus major vent. The expression 1 - (P-d/P-s) -
the delivery coefficient - where Pd is the dynamic pressure in the base of
the intact culm, and P-s the static pressure developed by the culm with the
outflow blocked, was identified as a useful and easily obtained measure of
the ability of an individual intact culm to contribute to the convective g
as-flow in a stand, and one which should be relatively unaffected by weathe
r conditions. For identical interlinked culms it was shown that the flows f
rom all culms fitted exactly along the same line of declining flow versus d
ynamic pressure irrespective of the numbers of culms or number and position
of outflow vents. (C) 2001 Elsevier Science B.V. All rights reserved.