Biocolonization and colour change, patina and microstromatolite format
ion were studied by analysing the mineralogy, chemistry, petrology and
biological activity on two granitic monuments in the Mediterranean ar
ea. Based on these analyses and laboratory experiments with isolated m
icro-flora, it is concluded that: i) black yeasts and meristematically
growing black fungi develop on and within granitic rocks and comprise
the same genera and species associated with both marble and limestone
monuments; ii) on sheltered but episodically wet surfaces, a surface
biofilm or microbial mat gives rise to mineral deposition which traps
air-borne dust resulting in a mono- or multilayered patina that can be
regarded as a micro-stromatolite or monument stromatolite; and iii) t
he main minerals in this patina are calcite, Ca-oxalate, gypsum and Ca
-phosphate, none of which is influenced by the substrate, and they mus
t thus be regarded as the product of the stromatolite forming micro-fl
ora.