Roots have a range of cell types that each contribute to the acquisition of
nutrients and their subsequent transfer to the xylem. The activities of th
ese cells must be coordinated to ensure that delivery of nutrients to the s
hoot occurs at a rate that matches the demands of growth. The partitioning
of transport processes between different cell types is thus essential for r
oots to function effectively. This partitioning in considered at the level
of proteins, organelles and cells in relation to the accepted concepts of h
ow nutrients are taken up by roots and delivered to the xylem. Using K+ as
an example, the evidence underpinning current concepts is examined, gaps in
understanding identified and the contribution of some new approaches asses
sed.